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Dr. Tim Nuttle

Dr. Tim Nuttle

Office: Weyandt 213A
Telephone: 724-357-2750

Office Hours

Visit Dr. Nuttle’s homepage.


B.S. Wildlife Ecology and Management – Michigan State University
M.S. Wildlife Ecology – Mississippi State University
Ph.D. Ecology – Utah State University


Forest ecology, restoration ecology, ecological modeling


Weyandt 213A Map

In the midst of the current Anthropocene mass extinction, loss of biological diversity seriously threatens ability of ecosystems to provide the services society relies on. Short of global extinctions at regional and local scales, biological communities are faced with many forces that threaten to simplify community structure. These impoverished communities may have reduced or altered function that can cascade across many trophic levels. Managers and decision-makers need to understand not only the effects of losses of biological diversity, but also their causal agents so that methods to manage, mitigate, and restore community structure and ecosystem function may be devised.

My research addresses the spectrum of this fundamental need: I seek to understand the processes by which diversity in biological communities develops, is maintained, and is lost, and the subsequent consequences of these changes on other ecosystem properties and processes. Please follow this link to read more about specific research projects being conducted in my lab



Temperton, V. M., R. J. Hobbs, T. Nuttle, and S. Halle. 2004. Assembly Rules and Restoration Ecology: Bridging the Gap Between Theory and Practice. Island Press, Washington, DC, USA.

Peer-reviewed Journal Articles

Anderson, JT, T Nuttle, JS Saldas Rojas, TH Pendergast, and AS Flecker. 2011. Extremely long-distance seed dispersal by an overfished Amazonian frugivore. Proceedings of the Royal Society of London B. doi:10.1098/rspb.2011.0155.

Nuttle, T., E. H. Yerger, S. H. Stoleson, and T. E. Ristau. 2011. Legacy of top-down herbivore pressure ricochets back up multiple trophic levels in forest canopies over 30 years. Ecosphere 2(1):art4. doi:10.1890/ES10-00108.1

Kansou, K., T. Nuttle, K. Farnsworth, and B. Bredeweg. 2011. Qualitative model of the long-term carbon cycle: the rise of the vascular plant during the Paleozoic period and its dramatic effect on our atmosphere. N. Agell and L. Rosello, eds. 25th International Workshop on Qualitative Reasoning, Barcelona, Spain, 16-18 July 2011.

Nuttle, T., and A. Bouwer. 2009. Supporting education about environmental sustainability: evaluation of a progressive learning route for qualitative models. Ecological Informatics 4:396-404.

Nuttle, T., B. Bredeweg, P. Salles, and M. Neumann. 2009. Representing and managing uncertainty in qualitative ecological models. Ecological Informatics 4:358-366.

Bredeweg, B., P. Salles, A. Bouwer, J. Liem, T. Nuttle, E. Cioaca, E. Nakova, R. Noble, A.L.R. Caldas, Y. Uzunov, E. Varadinova, and A. Zitek. 2007. Towards a structured approach to building qualitative reasoning models and simulations. Ecological Informatics 3: 1-12.

Nuttle, T., and J. W. Haefner. 2007. Design and validation of a spatially explicit simulation model for bottomland hardwood forests. Ecological Modelling 200:20-32.

Bredeweg, B., P. Salles, and T. Nuttle. 2007. Using exogenous quantities in qualitative models about environmental sustainability. AI [Artificial Intelligence] Communications 20(1):49-58.

Nuttle, T. 2007. Evaluation of restoration practice based on environmental filters. Restoration Ecology 15:330-333.

Leidolf, A., T. Nuttle, and M. L. Wolfe. 2007. Spatially scaled response of a lazuli bunting population to fire. Western North American Naturalist 67(1):1-7.

Nuttle, T., and J.W. Haefner. 2005. Seed dispersal in heterogeneous environments: bridging the gap between mechanistic dispersal and forest dynamics models. American Naturalist 165:336-349.

Nuttle, T., A. Leidolf, and L. W. Burger. 2003. Assessing the conservation value of bird communities with Partners in Flight–based ranks. Auk 120:541-549.

Leidolf, A., S. McDaniel, and T. Nuttle. 2002. The flora of Oktibbeha County, Mississippi. Sida 20(2):691-765.

Nuttle, T. 1997. Densiometer bias? Are we measuring the forest or the trees? Wildlife Society Bulletin 25(3):610-611.

Nuttle, T. and L. W. Burger. 1996. Response of breeding bird communities to restoration of hardwood bottomlands. Proceedings of the Annual Conference of Southeastern Fish and Wildlife Agencies 50:228-236

  • Biology Department
  • Weyandt Hall, Room 114
    975 Oakland Avenue
    Indiana, PA 15705-1081
  • Phone: 724-357-2352
  • Fax: 724-357-5524
  • Office Hours
  • Monday through Friday
  • 7:30 a.m.–12:00 p.m.
  • 1:00 p.m.–4:00 p.m.