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Dr. Sandra J. Newell

Dr. Sandy NewellContact Information
Office Hours


B.S., 1972, University of Toledo, Toledo, Ohio
Ph.D., 1976, University of Toledo, Toledo, Ohio
Postdoctorate, 1976-1978, Gray Herbarium, Harvard University


Population ecology, especially plants; evolutionary ecology and life history strategies; structure of clonal organisms; tropical ecology.


Weyandt 19 MapMy research interests are in plant population biology, plant-animal interactions, and carnivorous plants. My current research focuses on temperate and tropical pitcher plants. Videotaping is being used to determine the proficiency of insect capture by the plant. One population of pitcher plants was remarkably poor at capturing insects. Consequently, we have initiated a comparative study of several populations of one species in widely separated geographic locations (New Jersey, Michigan, and South Carolina). In addition, we are comparing the proficiency of insect capture in two different species of pitcher plant in South Carolina. Videotaping is also being used to analyze the behavior of insects, both prey and inhabitants, at the pitcher. My past research involved geographic variation in leaf morphology of cycads in Florida and Puerto Rico and demography in clonal violets.



NEWELL, S.J. and A.J. Nastase. 1998. "Efficiency of insect capture by Sarracenia purpurea (Sarraceniaceae), the northern pitcher plant," American Journal of Botany 85:88-91.

Newell, S.J. 1994. Occurrence of goldenrod galls: study of insect ovipositing behavior. Amer. Biol. Teacher 56(l):51-54.

Newell, S.J. 1989. Effects of position within a leaf on leaflet morphology in eight populations of Caribbean cycads ( Zamia pumila L.). Bull Torrey Bot. Club 116:229-239.

Newell, S.J. 1989. Variation in leaflet morphology among populations of Caribbean cycads ( Zamia). Amer. 1. Bot . 76:1518-1523.


Deppe, J. L., W. J. Dress, A. J. Nastase, S. J. Newell, C. S. LUCIANO. 2000. Diel variation of sugar amount in nectar from pitchers of Sarracenia purpurea L. with and without insect visitors. American Midland Naturalist 144: 123-132.

Dress, W.J., S.J. Newell, A.J. Nastase, and J.C. Ford. 1997. "Analysis of amino acids in nectar from pitchers of Sarracenia purpurea (Sarraceniaceae)," American Journal of Botany 84:1701-1706.

Nastase, A.J., C. De La Rosa, and S.J. Newell. 1995. Abundance of pitcher-plant mosquitoes, Wyroniyia smithii (Coq.) (Diptera: Culicidae), and midges, Metriocnemus knabi Coq. (Diptera: Chironomidae), in relation to pitcher characteristics of Sarracenia purpurea L. Amer. Midl. Nat ., 133:44-51.

Cipollini, D.F., Jr., S.J. Newell, and A.J. Nastase. 1994. Total carbohydrates in nectar of Sarracenia purpurea L. (northern pitcher plant). Amer. Midl. Nat . 131: 374-377.

Nastase, A.J., C. De La Rosa, and S.J. Newell. 1991. A comparison of three methods for collecting Dipteran insect larvae which inhabit the northern pitcher plant ( Sarracenia purpurea). Amer. Midl. Nat . 125:356-359.

Solbrig, O.T., W.F. Curtis, D.T. Kincaid, and S.J. Newell. 1988. Studies on the population biology of the genus Viola . VI. The demography of V. fimbriatula and V. lanceolata 1. Ecol. 76:301-319.

  • Biology Department
  • Weyandt Hall, Room 114
    975 Oakland Avenue
    Indiana, PA 15705-1081
  • Phone: 724-357-2352
  • Fax: 724-357-5524
  • Office Hours
  • Monday through Friday
  • 8:00 a.m.–12:00 p.m.
  • 12:30 p.m.–4:00 p.m.